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Department of Food Microbiology & Toxicology
UW-Madison


Spring 1999, Vol. 11, No. 1                    April 1999
Perspective
  • Faculty Positions in Food Safety
Research
Faculty & Staff
  • Agnes R. Denes, Ph.D., Research Associate
Short Subjects
  • Presentations
World Literature

 
Perspective
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The UW–Madison has released 32 new faculty positions in a campus-wide competition, with the goal of enhancing already existing high-priority campus programs. Based on FRI’s proposal, four of these new faculty positions have been consigned to food safety. The preliminary announcement follows:.

Faculty Positions in Food Safety

The University of Wisconsin–Madison has announced the development of a cluster of tenure-track faculty positions in food safety and will invite applications from qualified individuals. Successful candidates will develop innovative, responsive and highly competitive research programs that interact closely with other faculty at U.W.– Madison who conduct research in food safety. Successful candidates will have primary or joint faculty appointments as appropriate in participating departments where they will provide instructional and research leadership, and will coordinate their research/outreach activities through the Food Research Institute of the University of Wisconsin–Madison. Contact: Dr. Michael W. Pariza, Director, Food Research Institute, University of Wisconsin–Madison, 1925 Willow Drive, Madison, WI 53706.

Four positions are available for immediate recruitment, as follows:

  1. Emphasis on mycotoxins (70% research; 30% instruction)—The incumbent will investigate the biochemistry and genetics of mycotoxin synthesis by toxigenic molds, and the detection of mycotoxins in foods, feeds, and other biological materials, and provide undergraduate/graduate instruction in area(s) of expertise as appropriate.
  2. Emphasis on toxigenic molds (70% research; 30% instruction)—The incumbent will investigate the physiology and ecology of toxigenic molds, and factors that influence plant susceptibility and/or resistance to infection by toxigenic molds, and provide undergraduate/graduate instruction in area(s) of expertise as appropriate.
  3. Emphasis on immune response to foodborne pathogens and their toxins (70% research; 30% instruction)—The incumbent will investigate the host response to foodborne pathogens and their toxins, and the roles of foodborne pathogens and their toxins on immune modulation, and provide undergraduate/graduate instruction in area(s) of expertise as appropriate.
  4. Emphasis on novel engineering applications (70% research; 30% instruction)—The incumbent will investigate novel approaches to detecting and controlling foodborne pathogens and their toxins in foods, food processing equipment, and food manufacturing facilities, and provide undergraduate/graduate instruction in area(s) of expertise, for example in the food engineering/food science programs, as appropriate. Possible areas of specialty include surface modification, biosensors, and applicable instrumentation.


Research
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Comparative Survival of Salmonella typhimurium DT104, Listeria monocytogenes, and Escherichia coli O157:H7 in Preservative-free Apple Cider and Simulated Gastric Fluid  return to top

We compared the survival of three-strain mixtures (ca. 107 CFU/ml each) of Salmonella typhimurium DT104, Listeria monocytogenes, and Escherichia coli O157:H7 in pre-pasteurized and unpasteurized preservative-free apple cider (pH 3.3–3.5) during storage at 4 and 10°C for up to 21 d. Salmonella typhimurium DT104 populations decreased by <4.5 log10 CFU/ml during 14 d storage at 4 and 10°C in pasteurized cider, and by >5.5 log10 CFU/ml during 14 d in unpasteurized cider stored at these temperatures. However, after 7 d at 4°C, the S. typhimurium DT104 populations had decreased by only about 2.5 log10 CFU/ml in both pasteurized and unpasteurized cider. Listeria monocytogenes populations decreased below the plating detection limit (10 CFU/ml) within 2 d under all conditions tested. Survival of E. coli O157:H7 was similar to that of S. typhimurium DT104 in pasteurized cider at both 4 and 10°C over the 21 d storage period, but E. coli O157:H7 survived better (ca. 5.0 log10 CFU/ml decrease) than S. typhimurium DT104 (>7.0 log10 CFU/ml decrease) after 14 d at 4°C in unpasteurized cider. In related experiments, when incubated in simulated gastric fluid (pH 1.5) at 37°C, S. typhimurium DT104 and L. monocytogenes were eliminated (5.5–6.0 log10 CFU/ml decrease) within 5 and 30 min, respectively, whereas E. coli O157:H7 concentrations decreased only 1.60–2.80 log10 CFU/ml within 2 h. These results highlight the inherent differences in acid tolerance between these three pathogens and suggest that if S. typhimurium DT104, or even L. monocytogenes survive in apple cider, they are less likely than E. coli O157:H7 to survive passage through the stomach.

Ann M. Roering, John B. Luchansky, Anne M. Ihnot, Susan E. Ansay, Charles W. Kaspar, and S. C. Ingham


Recovery of Escherichia coli Biotype I during Refrigerated Storage of Beef Carcasses Inoculated with a Fecal Slurry
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The survival of Escherichia coli Biotype I on beef carcasses was investigated during a 10-day storage period at 4°C. Three beef carcasses were inoculated with a fresh slurry of cattle manure. Two sponge samples from each of 3 sites were taken from each of the three carcasses on days 0, 1, 3, 7, and 10 after inoculation and tested for E. coli Biotype I. The initial numbers ranged from an average of 171 cfu/cm2 to 405 cfu/cm2, depending on the counting method used (Petrifilm and MPN). In general, an appreciable reduction in the numbers occurred during the first 24 hours of storage, with an average reduction of 164 to 400 cfu/cm2, depending on the counting method. E. coli were not detected on day 7 of the storage period on 2 of the 3 carcasses, though viable E. coli were recovered from these 2 carcasses after a 24-hr enrichment at 37°C. Viable E. coli cells were detected on the third carcass after 7 days at 4°C. On day 10 viable cells were recovered from 2 of the 3 carcasses, but were not recovered from the third carcass. No significant difference in recovery of viable cells was observed between the two methods of enumeration on days 0, 1 and 3, though viable E. coli were recovered from all 3 carcasses on day 7. These findings suggest that i) refrigerated storage for more than 3 days may inactivate contaminating bacteria and improve safety, and ii) meat plants employing long term dry aging storage may test negative for E. coli, indicating such products are free of fecal contamination after 7 days, but such carcasses may still contain other bacterial indicators of fecal contamination, including enterococci.
— Mehmet Calicioglu, D. R. Buege, S. C. Ingham, and J. B. Luchansky


Growth and Biocontrol of Enterotoxigenic Bacillus cereus in Infant Formula
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We initiated a study to determine the potential for growth and enterotoxin production by Bacillus cereus in re-hydrated infant formula and to evaluate the effectiveness of various bacteriocins in controlling growth of the organism during storage at various temperatures. We inoculated 3 different rehydrated infant formulas (low iron and iron-fortified with and without maltodextrin) with 103 spores/ml of a 3-strain cocktail of B. cereus and stored the formulas at refrigeration (4, 8, and 12°C) and abuse (25°C) temperatures for up to 10 days. No growth occurred at any of the refrigeration temperatures. At 25°C growth began within 6 hours, and cell numbers reached 107 cfu/ml within 24 hr in all 3 formulations. Similar growth patterns occurred at lower inoculum levels (10 and 100 cfu/ml). When formula was inoculated with 103 spores/ml and stored at room temperature (25°C) for 6 hr before being refrigerated, outgrowth of spores occurred within 12 hr at 12°C and 8°C. Growth continued at 12°C over 48 hr, with cell counts reaching 105 cfu/ml. We found that 103 spores/ml of certain strains grew within 4 days in all 3 formulas at 25°C and 12°C. One strain grew at 8°C in the iron-fortified formula without maltodextrin, and another grew at 8°C in both of the iron-fortified formulas. We have developed an ELISA (enzyme-linked immunosorbent assay) that can detect as little as 0.25 ng/ml of each component of the tripartite B. cereus enterotoxin, hemolysin BL, and are assaying samples from the infant formula studies for the presence of toxin. So far we have observed that all 3 toxin components can be detected in formula after 24 hr of growth at 25°C. We are investigating the use of bacteriocins active against B. cereus. We tested five bacteriocins of lactic acid bacteria origin against a battery of B. cereus strains, using a spot-on-lawn in vitro assay. Results indicate that nisin is the most widely effective against the B. cereus strains tested, and is also the easiest to use, since it is commercially available in a standardized formulation. We have begun testing the efficacy of nisin in infant formulas against our 3-strain B. cereus cocktail and strain HRM44. Preliminary results show that 0.05% nisin (w/v) can inhibit the growth of strain HRM44 spores inoculated at 103 cfu/ml in formula at 8, 12, and 25°C for up to 10 days. Nisin also inhibits growth of the 3-strain cocktail at these temperatures, although the spores in the cocktail are less sensitive, and 0.10% nisin may be needed to provide inhibition. Results obtained to date indicate a potential for B. cereus to grow in re-hydrated infant formula during storage at 8°C or higher, and that the use of nisin can provide a hurdle to prevent outgrowth.
—Amy Wong and John Luchansky

 
Can Food Allergies Be Prevented or Cured?
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Food allergies affect 6–8% of children and 1–2% of adults in the United States. Reactions to foods can vary from mild episodes of hives to fatal anaphylaxis. Food allergic reactions are due to the development of IgE antibodies directed against specific proteins in the food. These antibodies circulate in the bloodstream and attach to the surface of mast cells in the tissues of the respiratory system, gastrointestinal system, and skin. When the food protein(s) come in contact with the IgE on the mast cell, histamine is released which leads to the allergic symptoms — hives, swelling in the skin, respiratory distress, vomiting, and shock. The propensity of an individual to produce IgE antibodies to food proteins is dependent upon genetic factors. Why certain foods, such as peanuts or fish, are more likely to cause allergy than other foods is unknown. Currently, the only ways to deal with food allergy are to avoid exposure, and administration of medications (epinephrin and antihistamines) when reactions occur. Since reactions can occur unexpectedly and with exposure to even small quantities of the food, new approaches are needed. Attempts to desensitize peanut allergic patients using conventional immunotherapy (“allergy shots”) have not been successful largely due to severe reactions to the treatment. Although improvements occurred in the threshold level of peanut exposure, reactions were not totally prevented. A new approach is the use of anti-IgE monoclonal antibodies which reduce the circulating levels of IgE. This treatment has been successfully used in the treatment of allergic rhinitis (“hay fever”) and allergic asthma by the Allergy Section at the University of Wisconsin–Madison. This treatment is far safer than traditional immunotherapy. Although this treatment has not been applied to food allergies, it will be investigated in the near future. Molecular biology studies of peanuts and other allergens indicate that certain amino acid sequences on the protein are necessary for the development of the IgE antibodies. Immunization with these peptides can shift the immune response away from the production of IgE. Molecular cloning of food allergens has led to the recognition that vaccination of animals with the DNA encoding for allergic proteins will also abrogate the IgE antibody response. Whether DNA-based vaccines can be used to prevent the development of allergy to specific food proteins has not been evaluated.
—Robert K. Bush, M.D.


Transgenic Alfalfa: A New System
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Countless examples of gene transfer have been published, but finding applicable products has been slow in development. A team of scientists with both basic and applied skills was organized to apply concepts of gene transfer to a working system of production. The team included Richard Burgess (former director of the Biotechnology Center), Sandra Austin-Phillips (gene transfer), Richard Straub (plant fractionation), Richard Koegel (harvest engineer), Edwin Bingham (alfalfa geneticist) and Mark Cook (animal scientist, FRI). Alfalfa was selected for gene transfer. UW–Madison has a long history of fractionation of juice proteins from alfalfa. In addition, alfalfa represents a significant crop raised in Wisconsin, and the unique color of alfalfa serves as a marker in a predominantly corn/soybean meal feeding system. Juices of alfalfa, expressing the desirable gene product, can be sprayed on foods post-pelleting, preserving gene product activity. Phytic acid is an inositol molecule that chelates plant phosphorus, rendering more than 60% of dietary phosphorus unavailable for digestion. Due to the unavailability of phytic phosphorus, animal diets must be supplemented with inorganic phosphorus which increases the cost of animal production in the US over $100 million per year. The enzyme phytase breaks down phytic acid and liberates the phosphorus, making it available for use in animal metabolism. It was theorized that the addition of phytase to animal feed would make phytic phosphorus available, thereby reducing the need to add inorganic phosphorus. Using phytic phosphorus would also decrease the levels of fecal phosphorus in animal excreta, and improve its value as a fertilizer. The phytase gene was successfully transferred from a fungus to alfalfa. The juice collected from alfalfa contained 1% of the soluble protein expressing phytase activity. When applied to chicken feed, the need for additional inorganic phosphorus was eliminated. Excreted phosphorus was reduced more than 50%. Seed stock is currently being generated in preparation for technology transfer. Studies on stability, fractionation, and animal performance continue. Other valuable genes are being inserted and expressed. This illustrates the ability of gene transfer to generate valuable products at a low cost. It is estimated that only 6% of the Wisconsin alfalfa crop will provide enough enzyme to meet the phytase needs of 7 billion broilers and 100 million pigs raised in the US. This illustration also demonstrates environmentally clean methods to advance the food production system.
—Mark E. Cook


Breakthrough in Transgenic Animal Technology
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Research reported in the 24 November 1998 edition of the Proceedings of the National Academy of Sciences (PNAS) may herald a new era in biopharmaceutical production. Work performed in the laboratory of former University of Wisconsin–Madison dairy science professor Robert Bremel, and by Gala Design LLC, a Sauk City, WI, company with which Bremel is now associated, resulted in new efficiency in the production of transgenic livestock. Dairy livestock are seen as key players in the production of genetically engineered protein drugs, such as monoclonal antibodies, hormones, vaccine proteins and enzymes. Introducing new genes into the animals allows them to produce specific pharmaceutical proteins in their milk, from where it can be extracted as a drug component. Costs of producing proteins by this route are far lower than traditional pharmaceutical production methods. The PNAS article describes a new method of gene introduction that increases the efficiency of production of transgenic cattle. The transgameticTM method inserts a gene into the unfertilized oocyte or egg, which stably incorporates the gene into the maternal germline. Once the egg is fertilized, all cells of the resulting embryo carry the new gene, and the calf is born with the capability to secrete a new protein in milk. Subsequent generations, offspring of each founder animal, will also carry the desired gene. Older production methods made transgenic livestock very costly. Cloning and pronuclear microinjection typically lead to only 1 percent of animals born carrying the new gene. The new technology also sidesteps problems of gene stability and mosaicism seen with microinjection. When DNA is microinjected into a fertilized embryo, the DNA is often not taken up until cell division has occurred. As a result, only some cell lineages carry the new gene. If the germ or sex cells don’t carry the new gene, the gene isn’t reliably transferred to offspring. This technology paves the way to applications of biotechnology in agricultural livestock, similar to those that have changed crop agriculture in recent years. The technology may also have applications in other mammals. The scientists believe the technology described also points to a key process in evolution. Transposable elements, or transposons, are known to jump to new locations in the genome (the cell’s hereditary material) and bring about changes in the genome that confer selective advantage. The experimental introduction of genes by a vector — which is structurally very similar to a transposable element — into the exposed genome of the oocyte results in uptake of new, fully heritable, genetic characteristics.
—UW–Madison Agricultural and Consumer Press Service

Faculty & Staff
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Agnes R. Denes, Ph.D., Research Associate 

 
Shortly after getting my B.S. in physical chemistry at the “Al. I. Cuza” University in Iasi, Romania, I came to the University of Wisconsin–Madison for my graduate studies. I received the Ph.D. in the Forestry Department in August 1998, working on cold-plasma-induced surface modification of natural and synthetic polymeric materials. I was involved in multiple research projects, one of them in collaboration with Dr. Amy Wong at the Food Research Institute. After finishing my graduate studies, I joined Dr. Wong’s research group. We are currently working on plasma-induced modification of organic and inorganic substrates to develop surfaces that inhibit biofilm deposition. 

Plasma-enhanced chemical processing is an excellent approach for surface modification of various biomaterials used in the food industry, and we hope that our work will bring us closer to creating materials that prevent the attachment of bacteria and the formation of biofilms. Our work also involves characterization of newly developed surface chemistries and properties using analytical tools available throughout the UW campus (e.g., electron spectroscopy for chemical analysis, atomic force microscopy, attenuated total reflection Fourier transform infrared spectroscopy). 

My husband is also in Madison, and he is a graduate student working on his Ph.D. in the Electrical Engineering Department. Aside from our common interest in the field of plasmas, the gaseous mixture of electrons, ions and neutral particles, we love traveling and the outdoors. When time permits, we are skating, swimming, playing tennis or walking our dog, Muki. Although we still miss the mountains from back home, we love Madison and its surroundings; we have met a lot of nice people and made good friends during our four years here.


Dr. Denes at electron spectroscope for chemical analysis.
Short Subjects
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Cook Presentations by Mark Cook:
“Conjugated linoleic acid on swine health and development.” Mark Cook at the North Carolina Swine Conference. Nov. 17, 1998. Raleigh, N.C. 
“Conjugated linoleic acid on poultry production.” Mark Cook at the North Carolina Poultry Conference. Dec. 16, 1998. Raleigh, N.C. 
Luchansky Presentations by John B. Luchansky:
“Practical examples of using PFGE to subtype bacteria from a variety of sources.” Invited speaker and participant at the CDC Workshop on Molecular Subtyping of Listeria monocytogenes and Escherichia coli O157:H7 by Pulsed-field Gel Electrophoresis. Centers for Disease Control and Prevention, Atlanta, Georgia. November 18, 1998. 

“Molecular tracking and biocontrol of bacteria associated with food.” Invited speaker at the Agricultural Research Service of the United States Department of Agriculture. The Eastern Regional Research Center, Wyndmoor, Pennsylvania. December 15, 1998. 

“Perceptions and prescriptions-Can Escherichia coli O157:H7 be controlled on farms or in foods?” Invited speaker at the 2nd Annual American Meat Science Association, National Meat Association Western Science Research Update. Doubletree Hotel and Conference Center, Monterey, California. February 17, 1999. 

Luchansky also traveled to Chicago in January to serve as the Technical Presentations Subcommittee Representative for the Biotechnology Division of the Institute of Food Technologists. 

In February, Luchansky and Senior Research Specialists Alan J. Degnan and Nan G. Faith hosted a group of undergraduate students for an afternoon at the FRI. This is the third year running that John, Al, and Nan gave students first-hand experience with genomic fingerprinting and biocontrol of E. coli O157:H7 associated with fermented meats. It was an enjoyable and productive experience for the mentors and the students.

Pariza Presentation by M. W. Pariza:
Co-organizer of the first “International Workshop on Conjugated Linoleic Acid Analysis,” held in Washington DC, January 1999.
 
World Literature
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Safety Concerns of Soy-based Infant Formula return to top
A recent article published by the Ministry of Health of New Zealand cautioned parents and medical professionals against the indiscriminate use of soy-based infant formulae. Isoflavone compounds, naturally present in soybean products, have been shown to adversely affect reproduction in some animal species and to have ill effects on some cells in culture. These compounds are weak estrogens and it has been suggested that they may have physiological effects on the neuroendocrine systems of infants who consume soy-based infant formulae.

Today, soy-based infant formulae account for approximately 25% of the formula sold in the U.S., and thousands of infants have consumed it with no overt signs of toxicity. Because of added mineral supplements which may contain incidental aluminum, soy formulae contain relatively high concentrations of aluminum (10–300 times the concentration of aluminum in human milk). Healthy full-term infants are not at substantial risk for aluminum toxicity but preterm and sick infants, particularly those with reduced renal function, may absorb excessive aluminum which can interfere with bone formation and nerve function.

Soy proteins are also allergenic. One of the main reasons that soy formula, rather than a cow’s milk-based formula, is fed to infants in the U.S. is the belief that it is less likely to provoke allergic reactions. However, a recent study of 263 Canadian infants, with a genetic predisposition to allergy, who were fed formulae either soy-based, cow’s milk-based or partial whey hydrolysate-based, compared to breast milk for at least the first 4 months, revealed that the incidence of asthma and eczema was similar in the infants fed the cow’s milk formula and the soy formula. In comparison, children who had consumed breast milk or the whey hydrolysate had about half the incidence of allergic reactions up to 5 years of age.

Phytoestrogens (isoflavones) are the latest safety concern associated with soy formulae. While some studies on carcinogenesis have demonstrated a protective effect of these compounds for certain cancers, other studies in animals and cell cultures have demonstrated deleterious effects. Plasma concentrations of isoflavones in soy-fed infants average 0.98 ug/ml compared to .005 ug/ml in infants on breast milk or cow’s milk formula. On a body weight basis, soy-fed infants consume about the same quantity of phytoestrogens as adults who consume moderate amounts of soy foods. However, Japanese adults who traditionally eat a variety of soy foods have plasma concentrations of isoflavones considerably lower than that measured in the soy-fed infants.

How biologically active are the plasma isoflavones? Isoflavones in soy-based infant foods are predominantly in the glucoside form which is unreactive. In the adult digestive tract, these conjugated forms can be readily hydrolyzed to form active compounds. However, we do not know how efficient infants are at hydrolysis of the glucosides.

Soy-based infant formulae, fortified with certain minerals and amino acids, have not been definitely implicated in any human illness or disorders with the exception of allergic reactions and aluminum toxicity in some preterm infants, but there does appear to be a potential for some adverse neuroendocrine effects caused by phytoestrogens. One concern is that these effects may not be manifested until many years later, for example, when the soy-fed infants reach puberty.

Reference: Food Chem News, Nov 9, 1998, p4.
—Ellin Doyle

Super-Potatoes?
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Since potatoes are a staple food in many countries, it has recently been proposed that varieties of potatoes be developed which could act as vehicles to induce immunity to food and waterborne pathogens. If this were technically feasible, it might be an effective means for vaccinating populations, particularly in underdeveloped areas, against epidemic diseases such as cholera. Potatoes could perhaps induce immunity to foodborne infections from Salmonella or E. coli. Several research groups are now working to achieve these goals.

The gene for a cholera toxin B subunit pentamer, which binds to cell surface receptors in the mucosa of the intestinal tract, has been introduced into potato tissues. When raw tuber tissue from these transgenic plants was fed to mice, it was found to induce both mucosal and systemic (serum) antibodies. When the mice were challenged with cholera toxin, there was a 60% reduction in the production of diarrheal fluid in the intestine. The antibodies induced by the transgenic potatoes in the mucosal tissue inhibited binding of the cholera toxin and significantly reduced symptoms of cholera. Since humans don’t normally eat raw potatoes, potatoes were cooked to determine whether cooking would destroy their ability to induce anti-cholera antibodies. About 50% of the immunogenic activity remained after cooking, hence the researchers suggested that future experiments might be aimed at finding a suitable fruit, consumed raw, as a foodborne vector for the vaccine.

In another series of experiments, the gene for the heat-labile enterotoxin subunit produced by E. coli was introduced into potato tissues. Mature tubers were harvested from plants grown from these transformed tissues and were found to produce this toxin subunit which binds to the intestinal mucosa. As with the previous experiments, mice fed these raw potatoes developed mucosal and serum antibodies which provided partial immunity to the effects of the E. coli enterotoxin. Further experiments with 14 brave and healthy human volunteers, fed 50–100 g raw transgenic potatoes containing an average of 0.75 mg of the E. coli enterotoxin subunit, demonstrated that humans also develop both mucosal and serum antibodies after eating these transgenic potatoes.

Fears over the safety of genetically engineered potatoes have been aroused in England by results of some toxicity tests in rats. In these experiments, genes coding for a lectin from snowdrop bulbs were introduced into potatoes in an attempt to make them more pest resistant. Some of the rats fed these potatoes exhibited adverse effects on their immune systems and on some of their organs, including kidney, spleen, and brain. These results have been disputed by other workers in the field and pending confirmation, should be considered as preliminary. Lectins are considered antinutritional for humans, though they are denatured or destroyed by heat, so cooked potatoes may be safe.

—Ellin Doyle

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Copyright © 1999 Food Research Institute
Last modified: 5 October 1999

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